Please visit my Google scholar page for an updated list of publications and citation information. I usually post preprints, but copyrights of published papers often belong to publishers. Please email me if you need a copy of one of my papers.

journal articles


  1. Cesar Nieto, Khem Raj Ghusinga, Cesar A. Vargas-Garcia, and Abhyudai Singh, “First-passage time statistics for size-dependent gene expression in growing cells: comparison of different models”, IEEE Control Systems Letters (under review), 2021.

  2. Khem Raj Ghusinga, Timothy C Elston, and Alan M Jones, “Towards resolution of a paradox in plant G-protein signaling”, Plant Physiology, 2021 [link].

  3. Khem Raj Ghusinga, Roger D Jones, Alan M Jones, and Timothy C Elston, “Molecular switch architecture determines response properties of signaling pathways”, Proceedings of the National Academy of Sciences, 2021 [link] [preprint].

  4. Khem Raj Ghusinga, Franco Pardes, Alan M Jones, and Alejandro Colaneri, “Reported differences in the flg22 response of the null mutation of AtRGS1 correlates with fixed genetic variation in the background of Col-0 isolates”, Plant Signaling & Behavior, 2021 [link].

  5. Khem Raj Ghusinga, Andrew Lamperski, and Abhyudai Singh, “Moment analysis of stochastic hybrid systems using semidefinite programming”, Automatica, 2020 [link] [preprint].

  6. Andrew Lamperski, Khem Raj Ghusinga, and Abhyudai Singh, “Analysis and control of stochastic systems using semidefinite programming over moments”, IEEE Transactions on Automatic Control, 2019 [link] [preprint].

  7. Cesar A. Vargas-Garcia, Khem Raj Ghusinga, and Abhyudai Singh, “Cell size control and gene-expression homeostasis in single cells”, Current Opinion in Systems Biology, 2018 [link].

  8. Khem Raj Ghusinga, Cesar A. Vargas-Garcia, Andrew Lamperski, and Abhyudai Singh, “Exact lower and upper bounds on stationary moments in stochastic biochemical systems”, Physical Biology, 2017 [link] [preprint].

  9. Saurabh Modi, Cesar A. Vargas-Garcia, Khem Raj Ghusinga, and Abhyudai Singh, “Analysis of noise mechanisms in cell size control”, Biophysical Journal, 2017 [link] [preprint].

  10. Khem Raj Ghusinga, John J. Dennehy, and Abhyudai Singh, “First-passage time approach to controlling noise in the timing of intracellular events”, Proceedings of the National Academy of Sciences, 2017 [link] [preprint].

  11. Khem Raj Ghusinga, Cesar A. Vargas-Garcia, and Abhyudai Singh, “A mechanistic stochastic framework for regulating bacterial cell division”, Scientific Reports, 2016 [link] [preprint].

  12. Vishwesh Kulkarni, Aditya Paranjape, Khem Raj Ghusinga, and Naira Hovakimyan, “Synthesis of robust tunable oscillators using mitogen activated protein kinase cascades”, Systems and Synthetic Biology, 2010 [link].

peer-reviewed conference proceedings


  1. Zahra Vahdat, Khem Raj Ghusinga, and Abhyudai Singh, “Comparing feedback strategies for minimizing noise in gene expression event timing”, Mediterranean Conference on Control and Automation, 2021 [link] [preprint].

  2. Khem Raj Ghusinga and Abhyudai Singh, “Controlling event timing precision in gene expression with dynamic external disturbances”, IEEE Conference on Decision and Control, 2020 [link] [preprint].

  3. Khem Raj Ghusinga and Abhyudai Singh, “Regulating gene expression to achieve temporal precision”, International Symposium on Mathematical Theory of Networks and Systems, 2020 [link] [preprint].

  4. Zikai Xu, Khem Raj Ghusinga, and Abhyudai Singh, “Noise analysis in biochemical complex formation from stochastically produced components”, American Control Conference, 2020 [link] [preprint].

  5. Khem Raj Ghusinga and Abhyudai Singh, “Analysis of stochastic timing of intracellular events with gene switching”, Indian Control Conference, 2019 [link] [preprint].

  6. Madeline Smith, Khem Raj Ghusinga, and Abhyudai Singh, “Comparison of feedback strategies for noise suppression in protein level”, American Control Conference, 2019 [link] [preprint].

  7. Khem Raj Ghusinga, Vaibhav Srivastava, and Abhyudai Singh, “Driving an Ornstein-Uhlenbeck process to desired first-passage time statistics”, European Control Conference, 2019 [link] [preprint].

  8. Khem Raj Ghusinga, Andrew Lamperski, and Abhyudai Singh, “Estimating stationary characteristic functions of stochastic systems via semidefinite programming”, European Control Conference, 2018 [link] [preprint].

  9. Khem Raj Ghusinga, Mohammad Soltani, Andrew Lamperski, Sairaj Dhople and Abhyudai Singh, “Approximate moment dynamics for polynomial and trigonometric stochastic systems”, IEEE Conference on Decision and Control, 2017 [link] [preprint].

  10. Ximing Chen, Masaki Ogura, Khem Raj Ghusinga, Abhyudai Singh and Victor M. Preciado, “Moment-Closures with Quality Guarantees using Semi-definite Programming: Application to Epidemics on Networks”, IEEE Conference on Decision and Control, 2017 [link].

  11. Khem Raj Ghusinga and Abhyudai Singh, “Effect of gene-expression bursting on stochastic timing of cellular events”, American Control Conference, 2017 [link] [preprint].

  12. Andrew Lamperski, Khem Raj Ghusinga, and Abhyudai Singh, “Stochastic optimal control using semidefinite programming for moment dynamics”, IEEE Conference on Decision and Control, 2016 [link] [preprint].

  13. Khem Raj Ghusinga and Abhyudai Singh, “Optimal regulation of protein degradation to schedule cellular events with precision”, American Control Conference, 2016 [link] [preprint].

  14. Khem Raj Ghusinga and Abhyudai Singh, “Theoretical predictions on the first-passage time for a gene expression model”, IEEE Conference on Decision and Control, 2015 [link].

  15. Khem Raj Ghusinga, Pak-Wing Fok, and Abhyudai Singh, “Optimal auto-regulation to minimize first-passage time variability in protein level”, American Control Conference, 2015 [link].

  16. Khem Raj Ghusinga and Abhyudai Singh, “First-passage time calculations for a gene expression model”, IEEE Conference on Decision and Control, 2014 [link] [preprint].

  17. Khem Raj Ghusinga and Abhyudai Singh, “First passage time stochasticity in a gene network with feedback regulation”, Northeastern Bioengineering Conference, 2014 [link].